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Fraxinus excelsior

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Fraxinus excelsior, known as the ash, or European ash or common ash to distinguish it from other types of ash, is a flowering plant species in the olive family Oleaceae. It is native throughout mainland Europe<ref name=Thomas/> east to the Caucasus and Alborz mountains. The northernmost location is in the Trondheimsfjord region of Norway.<ref name=rushforth>Rushforth, K. (1999). Trees of Britain and Europe. Collins Template:ISBN.</ref><ref name=dvf>Den virtuella floran: Fraxinus excelsior distribution</ref> The species is widely cultivated and reportedly naturalised in New Zealand and in scattered locales in the United States and Canada.<ref>Kew World Checklist of Selected Plant Families, Fraxinus excelsior</ref><ref>Biota of North America Program, Fraxinus excelsior</ref><ref>Altervista Flora of the United States and Canada, Fraxinus excelsior</ref>

Description

Male flowers
Seeds of Fraxinus excelsior, popularly known as "keys" or "helicopter seeds", are a type of fruit known as a samara

It is a large deciduous tree growing to Template:Convert (exceptionally to Template:Convert) tall with a trunk up to Template:Convert (exceptionally to Template:Convert) diameter, with a tall, narrow crown.<ref name=Thomas>Template:Cite journal</ref> The bark is smooth and pale grey on young trees, becoming thick and vertically fissured on old trees. The shoots are stout, greenish-grey, with jet black buds (which distinguish it from most other ash species, which have grey or brown buds). The leaves are opposite, Template:Convert long, pinnately compound, with 7–13 leaflets with coarsely serrated margins, elliptic to narrowly elliptic, Template:Convert long and Template:Convert broad and sessile on the leaf rachis.<ref name=Thomas/> There are no stipules.<ref name=Thomas/> These features distinguish ash from mountain ash (Sorbus aucuparia) in which the leaves are alternate with paired stipules. The leaves are often among the last to open in spring, and the first to fall in autumn if an early frost strikes; they have no marked autumn colour, often falling dull green. The flowers are borne in short panicles, open before the leaves, and have no perianth. The female flowers are somewhat longer than the male flowers, dark purple, without petals, and are wind-pollinated. Both male and female flowers can occur on the same tree, but it is more common to find all male and all female trees. A tree that is all male one year can produce female flowers the next, and similarly a female tree can become male. The fruit is a samara Template:Convert long and Template:Convert broad, often hanging in bunches through the winter;<ref>Kilbracken, J.1983. Larousse Easy way guide Trees. Larousse. Template:ISBN</ref> they are often called 'ash keys'.<ref name=rushforth/><ref name=afm>Mitchell, A. F. (1974). A Field Guide to the Trees of Britain and Northern Europe. Collins Template:ISBN</ref><ref name=afm1>Mitchell, A. F. (1982). The Trees of Britain and Northern Europe. Collins Template:ISBN</ref> If the fruit is gathered and planted when it is still green and not fully ripe, it will germinate straight away, however once the fruit is brown and fully ripe, it will not germinate until 18 months after sowing (i.e. not until two winters have passed).<ref>Forestry Commission/ Edlin, H. L. Broadleaves, HMSO, 1985, p36</ref>

European Ash rarely exceeds 250 years of age. However, there are numerous specimens estimated between 200 and 250 years old and there are a few over 250. The largest is in Clapton Court, England and is Template:Convert in girth. There are several examples over Template:Convert in Derbyshire alone.

Distribution

Fraxinus excelsior is native to Europe from northern Spain to Russia, and from southern Fennoscandia to northern Greece.<ref name=Thomas/> It is also considered native in southwestern Asia from northern Turkey east to the Caucasus and Alborz mountains. The northernmost location is in the Trondheimsfjord region of Norway.<ref name=rushforth/><ref name=dvf/> The species is widely cultivated and reportedly naturalized in New Zealand and in scattered locales in the United States and Canada including Nova Scotia, New Brunswick, Quebec, Massachusetts, Connecticut, New York, New Jersey, Maryland, Ontario, Ohio, Kentucky and British Columbia.Template:Citation needed

It is native throughout the British Isles, particularly on limestone, as in northern Scotland, where the most northerly native ashwood in the British Isles occurs on limestone at Rassal Ashwood.<ref name=Stiven/> It is widely planted elsewhere.<ref name=BRCBSBI>Template:Cite web</ref>

Ecology

Ash occurs on a wide range of soil types, but is particularly associated with basic soils on calcareous substrates. The most northerly ashwood in Britain is on limestone at Rassal, Wester Ross, latitude 57.4278 N.<ref name=snh>Template:Cite web</ref> Ash prefers moister soil types and is commonly limited by temperature and so not found at the higher colder altitudes in much of Europe, though in Iran, it may reach 2000 m asl. As a young seedling, it is shade tolerant, but older trees are light-demanding. It is an early-succession species and may well outcompete beech and oak, which are later-succession species.<ref>A review of European ash (Fraxinus excelsior L.): implications for silviculture Forestry, Dorota Dobrowolska1,*, Sebastian Hein, Anne Oosterbaan, Sven Wagner, Jo Clark and Jens Peter Skovsgaard (2011) doi: 10.1093/forestry/cpr001, 2011</ref>

F. excelsior mycorrhizae are of the internal arbuscular mycorrhizal type, in which the fungus grows within the tissues of the root and forms branched, tree-like structures within the cells of the root cortex. Unlike other Fraxinus species, F. excelsior does not form ectomycorrhizae.<ref name=Thomas/>

The Biological Records Centre of the UK records 111 species of insects and mites using ash as a food plant, of which 29 are specific to ash. A further six are specific to ash and its Oleaceae relative wild privet (Ligustrum vulgare).<ref name="″toller″">Template:Cite book</ref> A number of Lepidoptera species use the species as a food source.<ref>Template:Cite web</ref> One example of an ash-specific feeding moth is the centre-barred sallow (Atethmia centrago). The larvae burrow into the buds when newly hatched and later feed on the flowers and leaves.<ref>Template:Cite web</ref> A common moth which causes the browning of ash leaves, as well as garden privet and lilac, is Gracillaria syringella. The usually gregarious larvae form an epidermal gallery (i.e. feed within the leaf) which leads to a brown blotch with black frass. Later, two successive cones are formed by folding the tip of a leaf downwards.<ref>Template:Cite web</ref>

In Britain, 14 galls have been recorded on ash. The British Plant Gall Society defines a gall as "... an abnormal growth produced by a plant under the influence of another organism".<ref name="bpg">Template:Cite book</ref>

Ash dieback

Ash dieback is caused by the fungus Hymenoscyphus fraxineus which was previously known as Chalara fraxinea. Research into the genetics of the resistance of ash (Fraxinus excelsior) has shown that resistance does occur in European populations, but at least for the samples tested, it is neither common nor strong.<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref name="pliura2011">Template:Cite journal</ref>

Genome

The genome of Fraxinus excelsior is being sequenced by two groups of scientists in the United Kingdom. A group at Queen Mary University of London led by Richard Buggs are sequencing the self-pollinated offspring of a tree from Worcestershire, held by the Earth Trust.<ref>Template:Cite web</ref> A group at the John Innes Centre and The Genome Analysis Centre led by Allan Downie are sequencing "Tree 35" from Denmark, discovered by Erik Kjær, which has survived 8 years of ash dieback.<ref>Template:Cite web</ref>

Uses

Veneer of common ash wood
Replica of the body frame from the Volvo ÖV 4 car, made primarily from ash wood

The resilience and rapid growth made it an important resource for smallholders and farmers. It was probably the most versatile wood in the countryside with wide-ranging uses. Until World War II, the trees were often coppiced on a 10-year cycle to provide a sustainable source of timber for fuel and poles for building and woodworking.<ref name=mabey>Mabey, R. (1996). Flora Britannica. Sinclair-Stevenson Ltd Template:ISBN.</ref> The colour of the wood ranges from creamy white to light brown, and the heart wood may be a darker olive-brown. Ash timber is hard, tough and very hard-wearing, with a coarse, open grain and a density of 710 kg/m3.<ref>Ash. Niche Timbers. Accessed 19-08-2009.</ref> It lacks oak's natural resistance to decay, and is not as suitable for posts buried in the ground. Because of its high flexibility, shock resistance, and resistance to splitting, ash wood is the traditional material for bows, tool handles, especially for hammers and axes, tennis rackets, and snooker cue sticks,Template:Citation needed and it was extensively used in the construction of early aircraft. Ash was commonly used green for making chair frames which would be seated with another timber or with woven rush (e.g. those made by Philip Clissett, see also The English Regional Chair<ref>Template:Cite book</ref>). The parts were turned on a pole lathe or shaped with a drawknife. The practice essentially died out in the early 20th century, but has seen a revival in recent years.

Ash is an important constituent of wood pasture, a European management system in which open woodland provided shelter and forage for grazing animals.<ref name=Stiven>Template:Cite book</ref> Ash was coppiced and pollarded, often in hedgerows, and evidence in the form of some huge boles with multiple trunks emerging at head height can still be seen in parts of Britain. The Glen Lyon ash is a notable example of a pollarded ash which at about 400–500 years of age achieved a girth of 6 m.<ref name=Stiven/> In Northumberland, crab and lobster pots (traps) sometimes known as 'creeves' by local people are still made from ash sticks.Template:Citation needed Because of its elasticity European ash wood was commonly used for walking sticks. Poles were cut from a coppice and the ends heated in steam. The wood could then be bent in a curved vise to form the handle of the walking stick. The light colour and attractive grain of ash wood make it popular in modern furniture such as chairs, dining tables, doors, and other architectural features and wood flooring.

Ash is the only wood used for the manufacture of hurleys, referred to as hurls in parts of Leinster and known as a camán in Irish, the timber sticks used in the game of hurling in Ireland. Hurleys are manufactured from the butt log (bottom 1.5-m of the stem) and from trees ideally of a diameter at breast height around 25–30 cm. Only fast-grown, straight, and knot-free ash can be used for this purpose. Due to the lack of available ash in Ireland, over 75% of the timber needed to produce the 350,000 hurleys required for the game annually must be imported, mostly from Eastern European countries.<ref>Template:Cite web</ref> The importance of ash timber to the game of hurling is reflected in the fact that the game is referred to all over Ireland as "The Clash of the Ash".

Ash is valuable as firewood because it burns well even when 'green' (freshly cut).<ref>http://www.scoutbase.org.uk/library/hqdocs/facts/pdfs/fs315001.pdf</ref> Ash bark and leaves are used in modern herbal medicine for its astringent and laxative properties.<ref>Template:Cite journal</ref>

Mythology

In the 13th-century Edda and other writing relating to Norse mythology, a mythological ash tree called Yggdrasil serves as the center of the world.

Folklore

On the Isle of Bute in Scotland, lovers reportedly used to eat leaves of an ash tree known as the "Dreamin' Tree" that grew near the church of St Blane, and the pleasant dreams they then experienced revealed their actual spouses and intended fates.<ref>Downie, R. Angus (1934). Bute and the Cumbraes Glasgow & London : BlacKwood & Son Ltd. Page 83</ref>

Cultivars

Ash saplings from a mast year

Its many cultivars include;

  • Fraxinus excelsior 'Aurea', the traditional, slow-growing golden ash- not to be confused with 'Jaspidea'<ref>http://www.kew.org/plants-fungi/Fraxinus-excelsior.htm</ref>
  • Fraxinus excelsior 'Aurea Pendula' (weeping golden ash)
  • Fraxinus excelsior 'Autumn Blaze'
  • Fraxinus excelsior 'Autumn Purple'
  • Fraxinus excelsior 'Crispa'
  • Fraxinus excelsior 'Diversifolia' (one-leaved ash)
  • Fraxinus excelsior 'Erosa'
  • Fraxinus excelsior 'Jaspidea' a modern, vigorous golden ash
  • Fraxinus excelsior 'Monophylla'
  • Fraxinus excelsior 'Nana'
  • Fraxinus excelsior 'Pendula' (weeping ash), one of the best-known cultivars, widely planted during the Victorian era, grows vigorously forming an attractive small to medium-sized tree with mounds of weeping branches
  • Fraxinus excelsior 'Skyline'

Gallery

References

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External links

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